Marshall_etal_2000.html
Marshall, Helen L., Richard J. Geider and Kevin J. Flynn 2000 A mechanistic model of photoinhibition photoinhibit New Phytol 145 347-359.
model simulates time dependent decrease in alpha due to D1 damage and non-photochemical quenching
damage to the D1 protein in PSII is a linear function of light dose <my comment - refer to poisson model>
D1 is damaged then actively repaired, turnover leads to steady state of photosynthesis rate in constant light
example of non-steady state proportion of D1 (figure 2) goes from 1 at low light, to .3 at 0.25 mol photons m^-2 s^-1 <my comment - which equals .25 /(3.6 ^3) = 70 umol photons m^-2 s^-1.
< my comment> it's hard to tease out the damage rate and repair rates from their equations
fluctuating light may lead to significantly higher integrated production because the cells can capture high light and then do repair under lower light conditions