5 March 2003

Final exam 19 March .... please attend

problem with final exam and course grade

several students each term have decrease in exam scores:

99 - 92 - 73

94 - 72 - 88

99 - 99 - 95


 Chapter 14 Cloning and manipulation of DNA

overview...... LibraryOverview.gif

 

restriction enzymes

Palindromes ..(not on web)... palindromes.gif

RestrictionEnzymes.GIF

and methylation  

vectors

"Bacterial plasmids are circular extrachromosomal elements that replicate autonomously within cells.."

"Plasmid cloning vectors are derivatives of these natural plasmids 'engineered' to have features useful for cloning DNA"

vectors and genome sizes

genomeandvector.GIF

the idea..From workbook page 42

Plasmid Foreign Sequence .... PlasmidForeignSeries.html

PlasmidForeign1.GIF

PlasmidForeign2.GIF

PlasmidForeign2a.GIF

PlasmidForeign3.GIF

Items needed for good vector (text p. 302)

a) origin sequence for replication in E. coli

b) unique restriction enzyme sites

c) selectable markers to identify which E. coli colonies contain a vector and which E. coli colonies with a vector have a vector with a DNA insert

Vector used in the text is pUC19

Picture of pUC19..... plasmidpUC19.gif

which plasmid has foreign DNA?...plasmid1.gif

which bacteria have the plasmid? ...plasmid2.gif

which colony has the DNA we are looking for? ......plasmid3.gif

Review of insertion of foreign DNA into vector

Figure 14.5..(not on web).. 14x5.gif

 Lambda vector ....... l

Figure 14.6 ... 14x6.gif

LambdaVector.gif

YAC - Yeast artificial chromosome

Figure 14.7 .. 14x7.gif

 BAC - Bacterial artificial chromosome

Use of modified F plasmid in E. coli


7 March 2003

 

 

review of illustrations in the text

insertion of foreign DNA into plasmid .... 14x5.gif

insertion of foreign DNA into lambda vector .... 14x6.gif

structure of yeast artificial chromosome (YAC) ....14x7.gif

construction of cDNA ... for library or probe .... 14x8.gif

attachment of sticky end to cDNA for cloning .... 14x9.gif

screening plasmid genomic library for specific DNA ..... 14x11a.gif .....14x11b.gif

construction of restriction map .... 14x13.gif

Southern blot to find DNA matching a cDNA probe .... 14x14.gif

SouthernBlot.gif

example (not on web) .... hbs.gif

 

Libraries

Genomic libraries - entire genome -for 3,000,000 - Kb human genome, need 920,000 clones in lambda

Chromosome libraries - sort chromosomes, use digest of only one chromosome

cDNA libraries - use reverse transcriptase to get DNA complementary to mRNA

 

Screening a library

Concept of the probe .... ProbeTarget.html

Determination of which colony has the DNA wanted see Figure 14.11

14x11a.gif; .... 14x11b.gif

 

DNA sequencing

Gilbert and Sanger Nobel Prize in Chemistry 1980

Sanger method discussed in text.

Use of dideoxynucleotide DNA precursor. Figures 14.16 and 9.5... 9.5; 14.16.gif

Technique in Figure 14.15..... 14x15.gif

Step by step of Sanger technique:

As a series of slides ......SangerSeqsSeries.html

 

Denature DNA, add primer.....SangerSequencing1.gif

plus dideoxy / adenine..............SangerSequencing2.gif

plus dideoxy / guanine .............SangerSequencing3.gif

plus dideoxy / cytosine ............SangerSequencing4.gif

plus dideoxy/ thymine ............SangerSequencing5.gif

all together on a gel ..............SangerSequencing6.gif

 

Polymerase Chain Reaction (PCR)

Wonderful method of in vitro DNA amplification: polymerase chain reaction

ways of making lots of a specific sequence of DNA

the ultimate: PCR

1993 Nobel Prize Chemistry

Picture... Mullus-1993-1.gif

also .... Mullis.gif

reference: Kary Mullis article in Scientific American April 1990

"....revelation came to me one Friday night in April, 1983, as I gripped the steering wheel of my car and snaked along a moonlit mountain road into northern California's redwood country." .....p56

"You're not going to believe this," I crowed. "It's incredible."

Lederberg comment: --- anyone could have thought of this

Mullis response

to make lots of DNA....from article (not on web)

 


10 March 2003

last week, just before final exam, a time of great stress.....

lambda vector 600,000 library human

YAC vector 12,000 library

 

Review PCR and continue

Sequence of temperatures and events as in Text Figure 14.19 .... PCRasText2.gif

Graph fixed and variable 3' ends ..... PCRFixedVariable.gif

Simple PCR movie...... PCRmovie.html

PCR made possible by use of a temperature resistant DNA polymerase: from the bacterium Thermus aquaticus , Taq polymerase

functions at a high temperature, not denatured by temperature of denaturation (72º C is optimum temp)

Thermal cycler..(not on web). thermalcycler.gif

 

The process is patented. Here are some statements from a recent Carolina Biological Company catalog: "Practice of the patented Polymerase Chain Reaction (PCR) requires a license. The licensing covers both thermal cyclers and reagents. Manufacturers of thermal cyclers or reagents may have their products licensed for use in PCR by paying licensing fees to Hoffman-LaRoche, Inc. and Perkin Elmer Corporation, who own the patent rights to PCR."

Kinds of DNA amplification

cloning - large pieces, slow, libraries - use of vectors: plasmids, phage, BAC, YAC

PCR - fast, needs primers, small pieces of DNA

 

The use of PCR and cloning in forensic science, anthropology, studies of evolution

genomic and mitochondrial DNA from Romanov bones ... http://users.rcn.com/web-czar/dna.htm

and, New Yorker magazine 21/28 Aug 95

Some ideas/techniques

hybridization DNA/DNA and DNA/RNA; the power of base pairing!

denaturation of DNA to single strands

use of probes to identify large pieces of DNA

restriction enzymes to cut DNA at specific sites

vectors - plasmids, phage, artificial chromosomes of yeast and bacteria

introduction of foreign DNA into vectors (transformation for bacteria)

transfer of bacteria/DNA to nitrocellulose filter

replicate plating - make a copy of a pattern of bacterial colonies

in vitro synthesis of DNA to make primers

electrophoresis and Southern blotting

 

 

And, to chapter 15... Chapter15.html