BIOGEOGRAPHY
OF THE PACIFIC NORTHWEST
Instructor:
Dr. Keith Hadley
Office: 424 M CH
Office Hours: MWF 11:30-12:30 or by appointment
Phone: 725-3078
E-mail:
hadleyk@geog.pdx.edu (Subject: Geog413)
Web
Site: http://web.pdx.edu/~hadley
COURSE
OBJECTIVE:
This course examines the biogeography of the Pacific Northwest from a historical
and ecological perspective. Major
topics include historical and current plant and animal distributions, the
abiotic constraints to species distributions, ecological communities, and the
exogenous (external) and endogenous (internal) processes that shape these
communities. Much of the course will
focus on ecological processes such as succession, disturbance, competition, and
coexistence. Biogeographical theory and
its application to management and human modification of ecological communities
will also be examined.
PREREQUISITES:
Introductory
biogeography, ecology, or consent of instructor.
OPTIONAL
TEXT:
Natural
Vegetation of Oregon and Washington.
Franklin, J.F. and Dyrness, C.T. 1988.
Oregon
State University Press, Corvallis, Oregon.
GRADING:
Final
grades will be based on:
(1) Class participation
(2) Article Reviews
(3) Take Home Final Exam
(4) Term Project
(5) Field Trip Participation and Summary
(6) Graduate Student Presentations
The grading for this course focuses on: 1) in
class student participation based on their reviews of the course readings, 2) a
term project, 3) field trip participation, and 4) a take home Final Exam. Course readings will focus on classic papers
that examine various biogeographic patterns and processes found in the Pacific
Northwest. The term project will allow
students to examine a particular aspect of the biogeography of the Pacific
Northwest in greater detail (see term project handout). For example, students may wish to examine a
particular biogeographic pattern such as the distribution of a particular
organism or community or a particular process such as succession, disturbance,
invasion, etc. The objective of the
Final Exam is to have the student to synthesize the course material in a
fashion that is personally meaningful.
Specifically, the Final Exam will consist of several questions the
student may choose from that will have an applied or theoretical slant such as
preserve management, the preservation of biodiversity, or the adaptive
characteristics certain communities have to a particular type of disturbance.
The
class will have 1-3 field trips towards the end of the term. These will most likely take place on
Saturdays and each will replace one of our class meetings. Each field trip will have a set of required
readings. Graduate students may choose
to lead portions of one trip in lieu of their class presentation.
Final
grades for the course will be curved.
Students who feel they have a learning disability that may affect their
performance are encouraged to meet with me as soon as possible so that
accommodations can be made.
The
following is a course outline covering the major course topics and reading
assignments. This is a tentative
schedule that may be adjusted as the term progresses. In addition to textbook assignments, students will be responsible
for weekly written reviews of the
reserved readings (see below) and for the material included on course
handouts. Reserved readings and copies
of the overhead masters referred to during lecture are available in the
Geography Department Office. You may
check out these materials for not more than two hours; after 4:45 PM they may
be checked out overnight but must be returned by 9 AM the following
morning.
TENTATIVE CLASS SCHEDULE¹
TOPIC DESCRIPTION
1 Environmental
Setting
2 History
and Development of PNW Vegetation
3 Current
Vegetation of the Pacific Northwest
4 Community
Classification and Analysis
5 Ecosystem
Processes: Succession
6 Disturbance
Ecology
7 Environmental
Gradients
8 Boundaries,
Edges, and Ecotones
9 Meadows,
Balds, and Prairies
10 Applied
Biogeography and Landscape Ecology
¹
Required readings for each topic are listed below; Franklin, J.F. and Dyrness,
C.T. (1988) refers to your text.
Each student will create an annotated bibliography
for the required readings (except for readings from the text). The bibliography
will consist of: 1) a full reference for the article (as given below) and 2) an
article summary not to exceed one page.
Each article review must
be completed before the discussion of the article in class. All reviews will be graded and returned to
the student with comments the following week.
All reviews must be prepared using
a word processing program.
Daubenmire, R. 1969. Ecologic plant geography of the Pacific
Northwest. Madroño 20:111-128.
Franklin,
J.F. and Dyrness, C.T. 1988.
Introduction and Physical Setting.
Kruckeberg,
A.R. 1969. Soil diversity and the
distribution of plants, with examples from western North America. Madroño
20:129-154.
Delting, L. 1968. Historical Background of the Flora of the
Pacific Northwest. University of
Oregon Natural History Museum Bulletin 13.
Wolfe,
J.A. 1978. A Paleobotanical
Interpretation of Tertiary Climates in the Northern Hemisphere. American
Scientist 66:694-703.
² Graduate students
read each article; undergraduate students read either Delting (1968) or Wolfe (1978).
Delting,
L.E. 1953. Relict islands of xeric
flora west of the Cascade Mountains in Oregon.
Madroño
12:39-47.
Franklin,
J.F. 1988. Pacific Northwest
Forests. In: Barbour, M.G. and Billings, W.D. (eds.).
North American Terrestrial Vegetation. Cambridge University Press, New York.
Johannessen,
C., Davenport, W.A., Millet, A., and McWilliams. 1971. The vegetation of
the
Willamette Valley. Annals
of the Association of American Geographers 61:286-302.
Taylor,
D.W. 1977. Floristic Relationships along the Cascade-Sierran axis. American
Midland Naturalist 97:333-349.
Waring
and Franklin 1979. Evergreen coniferous
forests of the Pacific Northwest. Science
204:1380-1386.
Franklin,
J.F., Cromack, K. Jr., Denison, W., McKee, A., Maser, C., Sedell, J., Swanson,
F., and
Juday, G. 1981. Ecological characteristics of old-growth Douglas-fir
forests. USDA,
USFS General Technical Report
PNW-118. Portland, OR.
Franklin,
J.F. and Dyrness, C.T. 1988. Major
Vegetational Regions.
Minore,
D. (1979) Comparative Autecological Characteristics of Northwestern Tree Species:
A
Literature
Review. USDA
Forest Service Technical Report, PNW-87.
Wilson,
M.V., Hibbs, D.E., and Alverson, E.R. 1991.
Native plants, native ecosystems, and native landscapes. Kalmiopsis
13-17.
³Graduate students read
all; undergraduates all except Minore (1979)
Weaver,
H. 1959. Ecological changes in the
ponderosa pine forest of the Warm Springs Indian Reservation in Oregon. Journal
of Forestry 57:15-20.
Fastie,
C. 1995. Causes and ecosystem consequences of multiple pathways of primary
succession at Glacier Bay, Alaska. Ecology 76:1899-1916.
Disturbance Ecology4
Fire
Agee,
J. K. 1994. Fire and weather
disturbances in terrestrial ecosystems of the eastern
Cascades. USDA, USFS General Technical Report PNW -GTR-320. Portland, OR.
Boyd,
R. 1986. Strategies of Indian burning
in the Willamette Valley. Canadian Journal of Anthropology
5:65-86.
Kauffman,
J.B. 1990. Ecological Relationships of
vegetation and fire in Pacific Northwest
Forests. Pp.39-52. In:
Walstad, J.D., Radosevich, S.R., and Sandberg, D.V. (eds.).
Natural
and Prescribed Fire in Pacific Northwest Forests. Oregon State University Press.
Corvallis, Oregon.
Morris, W.G. 1934. Forest fires in Oregon and Washington. Oregon
Historical Quarterly 35:313-339.
4Graduate
students read all; undergraduate students read Boyd (1986), Kauffman (1990),
and Morris (1934)
Hadley,
K.S. and Savage, M.. 1996. Wind disturbance and development of a
near-edge forest interior, Marys Peak, Oregon Coast Range. Physical
Geography 17:47-61.
Taylor,
A.H. 1990. Disturbance and persistence
of Sitka spruce (Picea sitchensis
(Bong) Carr.) in coastal forests of the Pacific Northwest, North America. Journal
of Biogeography 17:47-58.
Insect Outbreaks
Speer,
J., Swetnam, T.W., Wickman, B.E., and Youngblood, A. 2001. Changes in Pandora
moth outbreak dynamics during the past 622
years. Ecology 82:679-697.
Grazing
Belsky, A.J. and Blumenthal, D.M.
1997. Effects of livestock grazing on
stand dynamics and soils in upland forest
of the interior West.
Conservation Biology 11:315-327.
Rummell,
R.S. 1951. Some effects of livestock
grazing on ponderosa pine forest and range in central Washington. Ecology
32:594-607.
Volcanic Eruptions
Halpren, C.B., Frenzen, P.M., Means, J.E.,
and Franklin, J.F. 1990. Plant
succession in areas of scorched and blown-down forest after the 1980 eruption
of Mount St. Helens, Washington. Journal of Vegetation Science 1:181-194.
Yamaguchi,
D. 1993. Forest History, Mount St. Helens. National Geographic
Research and Exploration 9:294-325.
Multiple Disturbance Agents
Geiszler,
D.R., Gara, R.I., Driver, C.H., Gallucci, V.F., and Martin, R.E. 1980. Fire, fungi, and
beetle
influences on a lodgepole pine ecosystem in south-central Oregon. Oecologia
46:239-243.
Parker,
A.J. 1995. Comparative gradient
structure and forest cover types in Lassen Volcanic and Yosemite National
Parks, California. Bulletin of the Torrey Botanical Club 122:58-68.
Zobel,
D.B., McKee, A., and Hawk, G.M. 1976.
Relationships of environment to composition, structure, and diversity of
forest communities of the central western Cascades of Oregon.
Ecological
Monographs 46:135-156.
Chen,
J., Franklin, J.F., and Spies, T.A. 1992.
Vegetation responses to edge environments in old-growth Douglas-fir
Forests. Ecological Applications 2:387-396.
Hadley,
K.S. 1999. Forest History and
Meadow Invasion at the Rigdon Meadows Archeological
Site, Western Cascades, Oregon. Physical Geography
20(2):116-133.
Magee,
T.K. and Antos, J.A. (1992) Tree
invasion into a mountain-top meadow in the Oregon
Coast Range, USA. Journal
of Vegetation Science 3:485-494.
Taylor, A.H. 1990. Tree invasion in meadow of the Lassen
Volcanic National Park. Professional Geographer 42:457-470.
Franklin,
J.F. and Forman, R.T.T. 1987. Creating landscape patterns by forest
cutting: Ecological consequences and principles. Landscape Ecology
1:5-18.
Orians,
G.H. 1990. “New Forestry” and the
old-growth Forests of Northwestern North America: a conversation with Jerry F.
Franklin. The Northwest Environmental Journal 6:445-461.
Zoobiogeography of the Pacific Northwest5
Ralph,
J.C., Hunt, G.L. Jr., Raphael, M.G., and Piatt, J.F. 1995. Ecology and Conservation of
the Marbled
Murrelet in North America: An Overview.
Chapter 1, Pp 3-22. In: Ralph, J.C.,
Hunt, G.L. Jr., Raphael, M.G., and Piatt, J.F. (Technical Editors),. Ecology and Conservation of the Marbled Murrelet. USDA, USFS, General Technical Report PSW-GTR-152. Albany, California.
Carey,
A.B., Horton, S.P., and Biswell, B.L. 1992.
Northern spotted owls: influence of prey base and landscape
character. Ecological Monographs 62:223-250.
Sanford,
E. 1999. Regulation of keystone
predation by small changes in ocean temperature.
Science
283:2095-2097.
5Read
either Ralph et al. 1995 or Carey et al. 1992.
Whittaker R. 1961. Vegetation history of the Pacific Coast States and the “central” significance of the Klamath Region. Madroño 16:5-23.